In which currency are paid payoffs in evolutionary games?

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In which currency are paid payoffs in evolutionary games? Krzysztof Argasinski Deparetment of Mathematics University of Sussex

Charles Darwin

Population growth equation (Malthus Law)

Payoff function which describe per capita growth rate  (number of new produced individuals) 

Evolutionary Game Theory

Evolutionarily stable strategies

John Maynard Smith

John von Neumann

Oskar Morgenstern

HAWK­DOVE GAME

Opponent

Player

(V-C)/2

V

0

V/2

gołąb 

V‐winners benefit C‐cost of conflict

C>V Hawk strategy dominate

C ≤V

Population growth equation

Payoff function which describe per capita growth rate  (number of new produced individuals) 

By following transition of coordinates:

We obtain following system of equations described on related frequencies Called…

Replicator dynamics:

Population growth equation

N(t) Payoff function which describe per capita growth rate  (number of new produced individuals) 

Problem with approach related to malthusian parameter with logistic suppression

Solution: turnover of individuals (Argasinski, Kozłowski, 2008)

Form of the nonlinear fitness function in classical evolutionary game theory…

n

W( p , pmean )   p Wj  pmean  i

j 1

i j

Economical application of game theory… Examples of mixed strategy phenotypes:

buy

P1

sell

buy

sell P2

Biological application of game theory…

Examples of mixed strategy phenotypes:

aggresive

P1

peacefull

aggresive

peacefull

P2

…so classic theory is focused on effects of elementary actions (pure strategies)

Problem of hidden assumptions

Problem of hidden assumptions Mathematical structure bias

Are those assumptions  testable?

Problem of hidden assumptions Mathematical structure bias

Are those assumptions  testable?

We  should compare  predictions of the model with empirical data  to show that there is something wrong

How can We measure parameters such costs and benefits?

How can We measure parameters such costs and benefits?

None of them will be pregnant in effect of this interaction

Therefore models related on malthusian parameter are not cause‐effect models.  What happens when we forget about malthusian parameter?

In effect we obtain generalization of replicator dynamics called 

SEX &VIOLENCE EQUATIONS

In effect we obtain generalization of replicator dynamics called 

SEX &VIOLENCE EQUATIONS

In effect we obtain generalization of replicator dynamics called 

SEX &VIOLENCE EQUATIONS

We can imagine case when mortality acts before reproduction or individuals  draw opponent that can kill them or mate (or both). This leads to variant called SEX OR VIOLENCE EQUATIONS

We can imagine more complicated cases, for example density dependent selection with two types of mortality pressures, one affecting fertility and second not:

When we apply new approach to the hawk dove game, we have two payoff matrices:

Survival matrix                                                   fertility matrix

When we apply new approach to the hawk dove game, we have two payoff matrices:

Survival matrix                                                   fertility matrix

New model is totally equivalent to classical Maynard‐Smith approach, however in new terminology we have:

B=W C=d(W+2)

In new model hawks may be stable

GAME OVER